I created a new email account (simhof@rosinto.com) on server #1 tonight and I tested inbound to the new email account on server #1 from Server #2. I get the following error (from tbird and roundcube) when attempting to send to the new email account (simhof@rosinto.com) from server #2. But interestingly, email from server #1 to server #1 works properly as local email. I tried tbird and roundcube both and got the same error.
An error occurred while sending mail. The mail server responded:
550 5.1.1 <simhof@rosinto.com>: Recipient address rejected: User unknown in virtual mailbox table.
Please check the message recipient "simhof@rosinto.com" and try again.
This is only happening to new email accounts on the server #1. Previously created email accounts on server #1 work fine.